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grinding of "@"?

In zwei in Release 5.0 [GC'd Beta Test rev 2], on Lisp Machine Janis Joplin (3600):

Why does having an embedded @ in the name make defun grind differently?

(defun foo ()
  bar)

(defun @ ()
       bar)

(defun -@ ()
  bar)

(defun @- ()
  bar)

(defun -@- ()
       bar)